1. Department of Cell and Developmental Biology; Graduate School of Biostudies; Kyoto University; Sakyo-ku Kyoto 606-8502 Japan
2. AMED-CREST; 1-7-1 Otemachi Chiyoda-ku Tokyo 100-0004 Japan
3. Institute for Integrated Cell-Material Sciences (WPI-iCeMS); Kyoto University; Sakyo-ku Kyoto 606-8507 Japan
4. Department of Life Science Frontiers; Center for iPS Cell Research and Application (CiRA); Kyoto University; Sakyo-ku Kyoto 606-8507 Japan