1. Wildlife Conservation Research Unit; Department of Zoology; The Recanati-Kaplan Centre; University of Oxford; Tubney House, Tubney; Abingdon; OX13 5QL; UK
2. Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Mengla; Yunnan; 666303; China
3. State Key Laboratory of Vegetation and Environmental Change; Institute of Botany; Chinese Academy of Sciences; No. 20 Nanxincun, Xiangshan; Beijing; 100093; China