1. Laboratory of Veterinary Hygiene, Department of Veterinary Medicine, Faculty of Agriculture, Tottori University, 4–101 Koyama Minami, Tottori 680–8553, Japan
2. Chiba Prefectural Livestock Research Center, He 16–1 Yachimata, Yachimata, Chiba 289–1113, Japan
3. Avian Zoonosis Research Center, Faculty of Agriculture, Tottori University, 4–101 Koyama Minami, Tottori 680–8553, Japan