1. Key Laboratory of National Forestry and Grassland Administration on Biodiversity Conservation in Karst Mountainous Areas of Southwest China, Guiyang, Guizhou 550001, China
2. Institute of Mountain Resources of Guizhou Province, Guizhou Academy of Sciences, Guiyang, Guizhou 550001, China
3. Guizhou Botanical Garden, Guizhou Academy of Sciences, Guiyang, Guizhou 550001, China