1. Animal Disease Control and Prevention Center in Qinghai Province, 69 Shengli Rd., Xining 810001, People's Republic of China;
2. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, 23 Xinning Rd., Xining 810001, People's Republic of China;
3. University of Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, People's Republic of China;
4. Department of Biology, The University of New Mexico, Albuquerque, New Mexico 87131, USA;