1. INRA UR0633, Zoologie Forestière, Orléans, France
2. Department of Entomology, Natural History Museum, London, UK
3. Botanischer Garten und Botanisches Museum Berlin‐Dahlem, Freie Universität Berlin, Germany
4. Biodiversity Institute of Ontario, University of Guelph, Ontario, Canada
5. Université de Rouen, ECODIV, UPRES‐EA 1293, Mont Saint Aignan, France
6. Department of Forestry, Technological Educational Institute of Kavala, Drama, Greece
7. contributed equally to the senior authorship