1. Institute of Comparative Medicine, Companion Animal Diagnostics, Bearsden Road, Bearsden, Glasgow, Scotland G61 1QH, UK
2. Centre for Integrated Genomic Medical Research, University of Manchester, Stopford Building, Oxford Road, Manchester M13 9PT, UK
3. Mammalian Immunogenetics Research Group, Faculty of Veterinary Science, University of Liverpool, Crown Street, Liverpool L69 7ZJ, UK
4. Department of Veterinary Pathology, University of Edinburgh, Easter Bush, Edinburgh, UK