1. 3Wellcome Sanger Institute, Hinxton, Cambridgeshire CB10 1SA, UK
2. 1London School of Hygiene and Tropical Medicine, Keppel Street, London WC1E 7HT, UK
3. 2Department of Medicine, University of Cambridge, Addenbrooke’s Hospital, Box 157 Hills Road, Cambridge CB2 0QQ, UK
4. 4Clinical Microbiology & Public Health Laboratory, Cambridge University Hospitals NHS Foundation Trust, Addenbrooke’s Hospital, Box 236 Hills Road, Cambridge CB2 0QW, UK