1. Quadram Institute Bioscience, Norwich, NR4 7UQ, UK
2. Department of Biosciences, Nottingham Trent University, Nottingham, NG11 8NS, UK
3. Faculty of Health and Life Sciences, University of Northumbria, Newcastle upon Tyne, NE1 8ST, UK
4. Norwich Medical School, University of East Anglia, Norwich, NR4 7TJ, UK