1. Tasmanian Institute of Agriculture, School of Land and Food, University of Tasmania, Sandy Bay, Hobart, Tasmania 7005, Australia
2. Faculty of Natural and Agricultural Sciences, University of the Free State, PO Box 339, Bloemfontein 9300, South Africa
3. Unité de Virologie et Immunologie Moléculaires, INRA Research Center, Domaine de Vilvert, 78352 Jouy-en-Josas cedex, France