1. 1European Centre for Environment and Human Health, University of Exeter Medical School, University of Exeter, Penryn, UK
2. 2Clinical Microbiology, Royal Cornwall Hospital, Truro, UK
3. 3Section for Food Safety and Emerging Health Threats, Norwegian Veterinary Institute, Oslo, Norway
4. 4Pathogen Informatics, The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, UK
5. †Present address: Quadram Institute Bioscience, Norwich Research Park, Norwich NR4 7UA, UK.