1. Illinois Natural History Survey, University of Illinois, 1816 South Oak Street, Champaign, IL 61820 USA
2. Graduate School of Agriculture, Tokyo University of Agriculture and Technology, 3-5-8 Saiwai-Cho, Fuchu, Tokyo 183-8509 Japan
3. Illinois Natural History Survey, University of Illinois, 1816 South Oak Street, Champaign, IL 61820 USA and Department of Natural Resources and Environmental Sciences, University of Illinois, 1102 South Goodwin, Urbana, IL 61801 USA