1. Division of Gastroenterology and Department of Medicine, Massachusetts General Hospital, Boston, Massachusetts 02114, USA;
2. Harvard Medical School, Harvard University, Boston, Massachusetts 02115, USA;
3. Novartis Institute for Biomedical Research, Cambridge, Massachusetts 02139, USA
4. Department of Cancer Immunology and Virology, Dana-Farber Cancer Institute, Boston, Massachusetts 02215, USA