1. Cancer Research UK London Research Institute, London, United Kingdom WC2A 3LY;
2. Present address: Translational Oncogenomics Lab, Centre for Evolution and Cancer, The Institute of Cancer Research, London, United Kingdom SW3 6JB
3. Centre for Mathematics & Physics in the Life Sciences & Experimental Biology (CoMPLEX), University College London, London, United Kingdom WC1E 6BT
4. Molecular and Population Genetics Laboratory, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom OX3 7BN;
5. Oxford National Institute for Health Research (NIHR) Comprehensive Biomedical Research Centre, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom OX3 7BN
6. University College London Hospital and Cancer Institute, CRUK Lung Cancer Centre of Excellence, London, United Kingdom WC1E 6DD