1. Wellcome Sanger Institute, Hinxton, Cambridgeshire CB10 1SA, United Kingdom;,
2. Theory of Condensed Matter, Department of Physics, University of Cambridge, Cambridgeshire CB3 0HE, United Kingdom
3. European Molecular Biology Laboratory, European Bioinformatics Institute (EMBL-EBI), Hinxton, Cambridgeshire CB10 1SA, United Kingdom