1. Division of Pulmonary, Critical Care, Allergy and Sleep Medicine, University of California, San Francisco, California, USA
2. Department of Medicine, University of California, San Francisco, California, USA
3. Current affiliation: Department of Pediatrics, School of Medicine and Public Health, University of Wisconsin–Madison, Madison, Wisconsin, USA
4. Department of Microbiology and Immunology, University of California, San Francisco, California, USA;
5. Howard Hughes Medical Institute, University of California, San Francisco, California, USA