1. Department of Pathology and Immunology Washington University School of Medicine, 660 S. Euclid Ave, St. Louis, Missouri 63110;
2. The Howard Hughes Medical Institute, Washington University School of Medicine, 660 S. Euclid Ave, St. Louis, Missouri 63110;
3. Department of Immunology and Microbiology Stanford University, Palo Alto, Callifornia 94305
4. The Howard Hughes Medical Institute, Stanford University, Palo Alto, Callifornia 94305