1. 1Jepson Herbarium and Department of Integrative Biology, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720-2465, USA;, Email: bbaldwin@berkeley.edu
2. 2University and Jepson Herbaria, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720-2465, USA;, Email: sfawcett@berkeley.edu
3. 3School of Biological Sciences, University of Portsmouth, Portsmouth PO1 2DY, UK