1. Department of Molecular Biology and Genetics, Cornell University, Ithaca, NY 14853, USA.
2. Department of Biology, Queen's University, Kingston, Ontario, K7L 3N6 Canada.
3. Biology Department, University of Pennsylvania, Philadelphia, PA 19104–6018, USA.
4. Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, PA 19104, USA.
5. Banting and Best Department of Medical Research and Department of Molecular and Medical Genetics, University of Toronto, Toronto, Ontario, M5G 1L6 Canada.