1. Department of Biology, University of British Columbia–Okanagan Campus, Kelowna, BC V1V 1V7, Canada.
2. Department of Integrative Biology, University of Guelph, Guelph, ON N1G 2W1, Canada.
3. Fort Keogh Livestock and Range Research Laboratory, U.S. Department of Agriculture–Agricultural Research Service, 243 Fort Keogh Road, Miles City, MT 59301–4016, USA.
4. MPG Ranch, Missoula, MT 59801, USA.
5. Department of Ecosystem and Conservation Sciences, University of Montana, Missoula, MT 59812, USA.