1. Centre for Marine Studies, Australian Research Council Centre for Excellence in Reef Studies and the Coral Reef Targeted Research Project, , The University of Queensland, St Lucia, Queensland (QLD) 4072, Australia;
2. Department of Biological Sciences, Macquarie University, New South Wales 2109, Australia;
3. Australian Commonwealth Scientific and Industrial Research Organisation (CSIRO) Sustainable Ecosystems, Post Office Box 284, Canberra Australian Capital Territory (ACT) 2601, Australia;
4. Fenner School of Environment and Society, The Australian National University, Canberra, ACT 0200, Australia;
5. Integrative Biology, 1 University Station C0930, University of Texas, Austin, TX 78712, USA;
6. The Ecology Centre, Centre for Applied Environmental Decision Analysis, The University of Queensland, St Lucia, QLD 4072, Australia;
7. Department of Biology, University of York, Post Office Box 373, York YO10 5YW, UK;