1. Department of Entomology, Pennsylvania State University, University Park, PA 16802, USA.
2. Center for Infection and Immunity, Mailman School of Public Health, Columbia University, New York, NY10032, USA.
3. Center for Infectious Disease Dynamics, Department of Biology, Pennsylvania State University, Mueller Laboratory, University Park, PA 16802, USA.
4. Fogarty International Center, National Institutes of Health, Bethesda, MD 20892, USA.
5. Bee Research Laboratory, U. S. Department of Agriculture–Agricultural Research Service, Beltsville, MD 20705, USA.