1. Department of Ecology and Evolutionary Biology, 2019 Kraus Natural Science Building, University of Michigan, 830 North University Avenue, Ann Arbor, MI 48109-1048, USA.
2. Center for Infectious Disease Dynamics (CIDD), Department of Biology, 208 Mueller Lab, Eberly College of Science, The Pennsylvania State University (PSU), University Park, PA 16802, USA.
3. Fogarty International Center, National Institutes of Health, Bethesda, MD 20892, USA.