1. College of Resources and Environmental Sciences, China Agricultural University, Beijing 100193, China.
2. Agri-Environment Branch, Agri-Food and Biosciences Institute, Belfast BT9 5PX, UK.
3. Department of Soil Science, Rothamsted Research, Harpenden, Herts AL5 2JQ, UK.
4. Department of Biology, Stanford University, Stanford, CA 94305, USA.