1. The Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, MD 21205–2185, USA.
2. Howard Hughes Medical Institute, Johns Hopkins University School of Medicine, Baltimore, MD 21205–2185, USA.
3. Department of Biochemistry and Molecular Biophysics, Howard Hughes Medical Institute, Columbia University, New York, NY 10032, USA.
4. Institut National de la Santé et de la Recherche Médicale (INSERM), UMR623, Developmental Biology Institute of Marseille (IBDM), Marseille 13288, France.