1. University of California, Irvine 1 Department of Ecology and Evolutionary Biology, 321 Steinhaus Hall , , Irvine, CA 92617 , USA
2. Oklahoma State University 2 Department of Integrative Biology , , Stillwater, OK 74078 , USA
3. University of California, Riverside 3 Department of Evolution, Ecology and Organismal Biology , , 900 University Avenue, Riverside, CA 92521 , USA