1. The Francis Crick Institute 1 Neural Development Laboratory , , 1 Midland Road, London NW1 1AT , UK
2. Centre for Craniofacial and Regenerative Biology 2 , Faculty of Dentistry, Oral and Craniofacial Sciences , , London SE1 1UL , UK
3. King's College London 2 , Faculty of Dentistry, Oral and Craniofacial Sciences , , London SE1 1UL , UK