1. Utah State University 1 Department of Biology , , Logan, UT 94321 , USA
2. University of California, Berkeley 2 Department of Environmental Science, Policy, and Management , , Berkeley, CA 94720 , USA
3. University of California, Berkeley 3 Museum of Vertebrate Zoology , , Berkeley, CA 94720 , USA
4. San Francisco State University 4 Department of Biology , , San Francisco, CA 94132 , USA
5. University of California, Berkeley 5 Integrative Biology , , Berkeley, CA 94720 , USA