1. Program in Developmental Biology, Washington University School of Medicine, St Louis, MO 63110, USA
2. Department of Genetics, Washington University School of Medicine, St Louis, MO 63110, USA
3. Laboratories of Genetics and Molecular Biology, University of Wisconsin, 1525 Linden Drive, Madison, WI 53706, USA
4. Department of Biology, Widener University, Chester, PA 19013, USA