1. Crucible Laboratory, Institute for Ageing and Health, Newcastle University, Newcastle upon Tyne, NE4 6BE, UK
2. Henry Wellcome Biogerontology Laboratory and Centre for Integrated Systems Biology of Ageing and Nutrition, Institute for Ageing and Health, Newcastle University, Newcastle upon Tyne, NE4 6BE, UK
3. School of Clinical and Laboratory Sciences, Newcastle University, Newcastle upon Tyne, NE4 6BE, UK
4. Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, NE4 6BE, UK