1. Graduate School of Science, Kyoto University 1 Division of Biological Sciences , , Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto City, Kyoto Prefecture 606-8502 , Japan
2. Graduate School of Agricultural and Life Sciences, The University of Tokyo 2 , 1-1-1 Yayoi, Bunkyo-ku, Tokyo Metropolitan 113-8657 , Japan
3. Sado Island Center for Ecological Sustainability, Niigata University 3 Marine Biological Station , , 87 Tassha, Sado City, Niigata Prefecture 952-2135 , Japan