1. Harvard Medical School 1 Angiogenesis Laboratory, Department of Ophthalmology, Massachusetts Eye and Ear Infirmary , , Boston, MA 02114 , USA
2. Massachusetts General Hospital, Harvard Medical School 2 Department of Molecular Biology , , Boston, MA 02114 , USA
3. Massachusetts General Hospital and Harvard Medical School 3 Department of Pathology , , Boston, MA 02114 , USA
4. Institute for Systemic Inflammation Research, University of Lübeck 4 , Lübeck 23562 , Germany
5. Cincinnati Children's Hospital Medical Center and University of Cincinnati College of Medicine 5 Division of Immunobiology , , Cincinnati, OH 45267, USA