1. College of Life Sciences, Capital Normal University 1 Laboratory of Stem Cell Biology , , Beijing 100048 , China
2. National Institute of Biological Sciences 2 , Beijing 102206 , China
3. College of Life Sciences, Nankai University 3 , Tianjin 300071 , China
4. State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University 4 , Tai'an 271018 , China