1. Wellcome Trust Sanger Institute, Wellcome Genome Campus, Hinxton, Cambridge CB10 1SD, UK
2. Department of Paediatrics, University of Cambridge, Cambridge CB2 0QQ, UK
3. Institute of Genetic Medicine, Newcastle University, Newcastle upon Tyne NE1 3BZ, UK
4. Theory of Condensed Matter, Theory of Condensed Matter Group, Cavendish Laboratory/Department of Physics, University of Cambridge, 19 JJ Thomson Avenue, Cambridge CB3 0HE, UK
5. Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne NE2 4HH, UK
6. Department of Dermatology and NIHR Newcastle Biomedical Research Centre, Newcastle Hospitals NHS Foundation Trust, Newcastle upon Tyne NE2 4LP, UK