1. Imperial College London 1 Department of Life Sciences , , London SW7 2AZ , UK
2. Peter Medawar Building for Pathogen Research, University of Oxford 2 , Oxford OX1 3SY , UK
3. University of Oxford 3 Nuffield Department of Clinical Laboratory Science , , Oxford OX3 9DU , UK
4. Oxford Brookes University 4 Department of Biological and Medical Sciences , , Oxford OX3 0BP , UK
5. UMR 7245 Molécules de Communication et Adaptation des Micro-organismes, Muséum National d'Histoire Naturelle, Centre National de la Recherche Scientifique (CNRS) 5 , CP52, 61 rue Buffon, 75231 Paris Cedex 05 , France