1. University of Aberdeen, 1 School of Medicine, Medical Sciences and Nutrition, Institute of Medical Sciences , Foresterhill, Aberdeen AB25 2ZD , UK
2. Instituto de Neurociencias de Alicante (Consejo Superior de Investigaciones Científicas-Universidad Miguel Hernández, CSIC-UMH) 2 , Campus San Juan, Av. Ramón y Cajal s/n, Alicante 03550 , Spain
3. , University College London 3 UCL Institute of Ophthalmology , 11-43 Bath Street, London EC1V 9EL , UK
4. Jena University Hospital, 4 Institute for Pharmacology/Toxicology , Drackendorfer Str. 1, D-07747 Jena , Germany