1. Department of Molecular and Cellular Biology and CREST/JST, Institute for Frontier Medical Sciences, Kyoto University, Sakyo-ku, Kyoto 606-8397, Japan
2. Department of Cell Biology, Nagahama Institute of Bio-Science and Technology, Nagahama, Shiga 526-0829, Japan
3. Institute for Molecular Science of Medicine, Aichi Medical University, Nagakute, Aichi 480-1195, Japan