1. Institute of Reproductive and Developmental Biology, Imperial College London, Hammersmith Hospital, Du Cane Road, London W12 0NN, UK
2. Department of Histopathology, Charing Cross Hospital, Fulham Palace Road, London W6 8RF, UK
3. Department of Mathematics and Centre for Integrative Systems Biology at Imperial College (CISBIC), Imperial College London, London SW7 2AZ, UK