1. Department of Hematology, Hokkaido University Faculty of Medicine, Graduate School of Medicine and School of Medicine, Sapporo 060-8638, Japan
2. Department of Cell Biological Science, Hokkaido University Graduate School of Life Science, Faculty of Advanced Life Science, Sapporo 060-0810, Japan
3. Innate Immunity Laboratory, Hokkaido University Graduate School of Life Science, Sapporo 001-0021, Japan
4. Research Institute for Biomedical Sciences, Tokyo University of Science, Chiba 162-8601, Japan