1. Department of Anatomy and Cell Biology, College of Medicine, Chungnam National University, Daejeon 35015, Republic of Korea
2. Department of Medical Science, College of Medicine, Chungnam National University, Daejeon 35015, Republic of Korea
3. Department of Otolaryngology-Head and Neck Surgery, College of Medicine, Chungnam National University, Daejeon 35015, Republic of Korea
4. Korea Drosophila Resource Center, Gwangju Institute of Science and Technology, Gwangju 61005, Republic of Korea
5. School of Life Sciences, Gwangju Institute of Science and Technology, Gwangju 61005, Republic of Korea
6. Department of Biological Sciences, Korea Advanced Institute of Science & Technology, Daejeon 34141, Republic of Korea