1. Department of Biotechnology, Graduate School of Life Sciences, Ritsumeikan University, Kusatsu 525-8577, Japan
2. Department of Life Sciences, Graduate School of Arts and Sciences, The University of Tokyo, Tokyo 153-8902, Japan
3. Department of Marine Science and Technology, Faculty of Marine Science and Technology, Fukui Prefectural University, Obama 917-0003, Japan
4. Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto 606-8502, Japan