1. School of Biomedical Sciences, The University of Hong Kong, Li Ka Shing Faculty of Medicine, Hong Kong, China
2. Genome Regulation Laboratory, CAS Key Laboratory of Regenerative Biology, Joint School of Life Sciences, Guangzhou Institutes of Biomedicine and Health, Chinese Academy of Sciences, Guangzhou 510530, China
3. Guangzhou Medical University, Guangzhou 511436, China
4. School of Biomedical Sciences, The Chinese University of Hong Kong, Shatin, Hong Kong, China
5. Department of Biology, College of Arts & Sciences, University of Iowa, Iowa City, IA 52242
6. Department of Otolaryngology, College of Arts & Sciences, University of Iowa, Iowa City, IA 52242
7. Francis Crick Institute, London NW1 1AT, United Kingdom