1. Graduate School of Science and Technology, Nara Institute of Science and Technology, Nara, 630-0192 Japan
2. Faculty of Science and Engineering, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, 9747 AG Groningen, The Netherlands
3. Department of Plant Biotechnology and Bioinformatics, Ghent University, 9052 Ghent, Belgium
4. VIB Center for Plant Systems Biology, 9052 Ghent, Belgium
5. Department of Developmental Biology, Institute for Plant Sciences and Microbiology, University of Hamburg, 22609 Hamburg, Germany