1. These authors contributed equally to this manuscript
2. National Poultry Laboratory Animal Resource Center, Harbin Veterinary Research Institute, CAAS, Harbin, China
3. State Key Laboratory for Animal Disease Control and Prevention, Harbin Veterinary Research Institute, CAAS, Harbin, China
4. Western Research Institute, CAAS, Changji, China
5. Department of Medical Biochemistry, University of Shizuoka School of Pharmaceutical Sciences, Shizuoka, Japan