1. Department of Applied Biological Resource Sciences, School of Agriculture, Ibaraki University, Ami-machi, Inashiki-gun, Ibaraki 300-0393, Japan
2. Department of Dermatology, School of Medicine, Kinki University, Osaka 589-8511, Japan
3. CNRS-University of Toulouse III UMR 5165, Epidermis Differentiation and Rheumatoid Autoimmunity, Institut Fédératif de Recherche 30 (INSERM, CNRS, CHU Toulouse-Purpan, Université Paul Sabatier), 37 allées Jules Guesde, 31073 Toulouse cedex 7, France