1. 1Leeds Institute of Rheumatic and Musculoskeletal Medicine (LIRMM), University of Leeds, Leeds, United Kingdom
2. 2National Institute for Health Research (NIHR) Leeds Biomedical Research Centre (BRC), The Leeds Teaching Hospitals, Leeds, United Kingdom
3. 3School of Molecular and Cellular Biology, University of Leeds, Leeds, United Kingdom
4. 4Department of Dermatology, University Medical Center of the Johannes Gutenberg University, Mainz, Germany