1. Department of Physiology and Biophysics, Weill Medical College of Cornell University, New York, New York, United States 2Burke Medical Research Institute, White Plains, New York, United States
2. Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States
3. Department of Ophthalmology and Visual Sciences, University of British Columbia, Vancouver, British Columbia, Canada
4. Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States 5Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States