1. Department of Ophthalmology, University of Washington, Seattle, Washington, United States
2. Evelyn F. and William L. McKnight Vision Research Center, Bascom Palmer Eye Institute, Department of Ophthalmology, University of Miami, Miami, Florida, United States
3. Department of Ophthalmology, University of Washington, Seattle, Washington, United States 3Departments of Biological Structure and Pathology, University of Washington, Seattle, Washington, United States