1. Department of Ophthalmology, University of California, San Francisco, San Francisco, California, United States
2. Department of Pathology, University of California, San Diego, La Jolla, California, United States
3. Advanced Scientific Research Leaders Development Unit, Gunma University, Gunma, Japan
4. Department of Ophthalmology, University of California, San Francisco, San Francisco, California, United States 4Department of Anatomy and Institute for Human Genetics, University of California, San Francisco, San Francisco, California, United States
5. Department of Pathology, University of California, San Diego, La Jolla, California, United States 5VA San Diego Healthcare System, San Diego, California, United States