1. Department of Neuroanatomy, Kanazawa University, School of Medicine, Kanazawa City, Ishikawa 920-8640, Japan
2. CREST, Japan Science and Technology Corporation, Kawaguchi, Saitama 332-0012, Japan
3. Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita City, Osaka 565-0871, Japan
4. Discovery Research Lab, TANABE SEIYAKU Co., Ltd., Osaka City, Osaka 532-0031, Japan
5. Skirball Institute, New York University School of Medicine, New York, NY 10016
6. Departments of Surgery, Physiology and Cellular Biophysics, College of Physicians and Surgeons of Columbia University, New York, NY 10032